Vascular plant communities of Spain and Portugal. Addenda to the Syntaxonomical checklist of 2001

Salvador Rivas-Martínez, Tomás E. Díaz, Federico Fernández-González, Jesús Izco, Javier Loidi, Mario Lousã & Ángel Penas

Itinera Geobotanica 15(1-2): 5-922 (2002)


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2. Syntaxon descriptions

Sambuco racemosae-Prunetum padi ass. nova hoc loco (66.5.1)

(Sambuco-Salicion capreae, Sambucetalia racemosae, Rhamno-Prunetea)

Typus associatio: Table 85, rel. 2 [Huesca: Benasque, Ball d'Alba. 42º 41'N-2º 35'E. 1710 m, NW, 20%, 60 m2].

Characteristic species (territorials): Lonicera alpigena, Prunus padus, Ribes rubrum, Sambucus racemosa.

Diagnosis: Upper supra and lower orotemperate humid and hyperhumid Pyrenean scrubby fast grow community dominated by Prunus padus, Salix caprea and Sambucus racemosa, growing on rich and wet soils in gullies and mountain cones as mantle of Abies alba or Pinus uncinata forests communities.

Table 85

66.5.1 Sambuco racemosae-Prunetum padi

(Sambuco-Salicion capreae, Sambucetalia racemosae, Rhamno-Prunetea)

Altitude (1=10 m)165171170175170
Number of species1610181414
Ordinal number12*345
Characteristic species:
Prunus padus23344
Sambucus racemosa12224
Rubus idaeus+12+4
Salix caprea.11+3
Ribes rubrum31.22
Rhamnus alpina2.3.2
Ribes alpinum..1.1
Companion species:
Lonicera alpigena11.33
Molopospermum peloponesiacum1.113
Dryopteris filix-mas.2113
Aconitum vulparia1.1 2
Sorbus aucuparia.1.+2
Corylus avellana..1+2

Other species: Companion species: Rosa pendulina 1 in 1, + in 4. Trisetum flavescens 1 in 2, 2 in 4. Chaerophyllum villarsii 1 in 2 and 4. Lilium martagon 2 in 3 and 4. Daphne mezereum +, Gentiana lutea +, Knautia arvernensis +, Phyteuma pyrenaicum +, Ranunculus platanifolius +, Trollius europaeus +, Veratrum album + in 1. Athyrium filix-femina 2, Peucedanum ostruthium 2, Ulmus glabra (S2) 2, Betula pubescens 1 (S2), Helleborus foetidus 1, Scrophularia alpestris 1, Sorbus aucuparia 1 in 3.

Localities: 1. Huesca: Benasque, Cono de avalanchas del Perdiguero. 50 m2. 2. Holotypus ass. Huesca: Benasque, Ball d'Alba. 42º 41'N-0º 35'E. NW, 20%, 60 m2. 3. Huesca: Benasque, Cono de avalanchas de la Tuca de Salvaguardia. W, 100 m2. 4. Lérida: Valle de Arán, Val de Molieres, Plan dera Rasa. 42º 37" 30"N-0º 45'E. S, 10%, 100 m2 5. Synthesized table.

[Rivas-Martínez, Costa & P. Soriano]

Saturejo salzmannii-Drosophylletum lusitanici (Galán & Vicente 1996) ass. nova, stat. nov. hoc loco (61.5.4)

[Stauracantho boivinii-Drosophylletum saturejetosum salzmannii Galán & Vicente in Bot. Helvetica 106: 51, tb. 1, rel. 39 to 43, in mistake rel. 22. 1996 (art. 5). New typus hoc loco, l. c., rel. 40.]
(Stauracanthion boivinii, Ulicetalia minoris, Calluno-Ulicetea)

Typus associatio: Holotypus hoc loco, Galán & Vicente in Bot. Helv. 106: 50, tb. 1, rel. 40. 1996.

Characteristic species (territorials): Drosophyllum lusitanicum, Satureja salzmannii.

Diagnosis: Association characterized by Drosophyllum lusitanicum and Satureja salzmannii growing on sandstone dystric-podzol leptosols in thermomediterranean humid bioclimatic belt of Aljibic biogeographic Sector (Coastal Lusitan-Andalusian Province); it is geovicariant of Stauracantho boivinii-Drosophylletum lusitanici Quézel, Barbero, Benabid, Loisel & Rivas-Martínez in Ecol. Medit. 14: 85, tb. 12. 1988, but it can be easely separated by Satureja salzmannii and the absence of Genista tridens or Pterospartum lasianthum and the dryer demands of Cisto-Lavanduletea species: Cistus crispus, Cistus monspeliensis, Lavandula stoechas subsp. atlantica, and Halimium halimifolium.

[Rivas-Martínez]

Saxifragetum bourgeanae ass. nova hoc loco (27.14.5)

(Campanulion mollis, Asplenietalia petrarchae, Asplenietea trichomanis)

Typus associatio: Málaga: Cortes de la Frontera, Sierra de Líbar. NE, 850 m, 10 m2. Characteristic species: 3 Saxifraga bourgeana, 3 Saxifraga granatensis, 2 Ceterach officinarum, 1 Sedum dasyphyllum, + Asplenium trichomanes subsp. quadrivalens, + Silene andryalifolia. Companion species: 1 Campanula specularioides, 2 Geranium purpureum, 1 Sedum album, 1 Umbilicus rupestris, + Arenaria arundana, + Biscutella frutescens, + Geranium lucidum, + Geranium rotundifolium, + Hedera helix, + Mucizonia hispida.

Characteristic species: Saxifraga bourgeana , Saxifraga granatensis.

Diagnosis: Chasmophytic calcareous and calcodolomitic community, growing in meso- and supramediterranean shady walls in the Rondean biogeographic Sector mountains.

[Díez Garretas, Asensi & Martín]

Saxifragetum retusae Gruber ass. nova hoc loco (27.6.8)

[Saxifragetum retusae Gruber 1978 nom. inval. (art. 1)]
(Androsacion vandellii, Androsacetalia vandellii, Asplenietea trichomanis)

Typus associatio: Andorra: Pic Siscarou. 42º31'N-1º33'E. N, 2600 m, 80%, 100 m2. Characteristic species: 2 Saxifraga retusa, 1 Saxifraga bryoides, + Asplenium septentrionale, + Cardamine resedifolia, + Draba dubia subsp. laevipes. Companion species: + Festuca airoides, + Minuartia recurva, + Saxifraga geranioides [relevé: M. Gruber 1978, tb. 1, rel. 1].

Characteristic species (territorial): Saxifraga retusa.

Diagnosis: Cryorotemperate hyperhumid Eastern Pyrenean chasmophyte calcifugous community, characterized by the alpine-pyrenean endemics Saxifraga retusa and Saxifraga bryoides, as well as the pyrenean endemic Saxifraga pentadactylis subsp. pentadactylis, growing on siliceous north exposed snow free walls.

[Rivas-Martínez]

Scrophulario auriculatae-Epilobietum hirsuti ass. nova hoc loco (40.5.6)

[Scrophulario auriculatae-Epilobietum hirsuti Ríos & Alcaraz in Ríos 1996 nom. inval. (art. 1)]
(Calystegion sepium, Calystegietalia sepium, Galio-Urticetea)

Typus associatio: Table 86, rel. 2 [Jaén: Santiago de la Espada, Huelga-Utrera. 30S WH3623. 1050 m, 40 m2].

Characteristic species (territorials): Epilobium hirsutum, Scrophularia auriculata, Scrophularia scorodonia, Solanum dulcamara, Eupatorium cannabinum, Calepina irregularis.

Diagnosis: Herbaceous forb community with Epilobium hirsutum, Eupatorium cannabinum, Scrophularia auriculata, Scrophularia scorodonia and Solanum dulcamara. It appears in river bank sediments rich in organic remains. It is well characterized by Scrophularia auriculata (= S. lyrata), species absent in the related associations. This community seems to be widespread in the West Iberian territories.

Table 86

40.5.6 Scrophulario auriculatae-Epilobietum hirsuti
(Calystegion sepium, Calystegietalia sepium, Galio-Urticetea)

Altitude (1=10m)1001051307898132978595120102
Number of species96109813117131210
Ordinal number12*3456789108
Characteristic species:
Epilobium hirsutum3554525245V
Scrophularia auriculata.+...213++IV
Eupatorium cannabinum21+.......III
Urtica dioica1.1......1III
Scrophularia scorodonia.+..+.....III
Solanum dulcamara..+.+.....II
Lycopus europaeus...1..+...II
Hypericum caprifolium.....1..+.II
Calystegia sepium....+.....I
Companion species:
Mentha longifolia+++..2..22V
Sonchus aquatilis+..+..+.+.III
Peucedanum hispanicum.+1+....+.III
Nasturtium officinale..+.+.+..+III
Ranunculus granatensis1.+......1III
Pulicaria dysenterica...++...2.III
Ranunculus repens...2...1+.III
Veronica beccabunga.....++..+III

Other species. Companion species: Hypericum undulatum + in 1 and 3. Equisetum ramosissimum + in 4 and 7. Lythrum salicaria + in 5, 1 in 8. Dorycnium rectum + in 5 and 8. Agrostis stolonifera + in 7 and 9. Calepina irregularis 2, Geranium robertianum + in 1. Lactuca virosa + in 3. Althaea officinalis +, Carex hispida + in 4. Cirsium ferox +, Epilobium parviflorum +, Equisetum palustre +, Hypericum tomentosum +, Juncus inflexus +, Lysimachia ephemerum +, Lysimachia vulgaris +, Veronica anagalloides + in 6. Equisetum arvense +, Juncus subnodulosus +, Scirpoides holoschoenus + in 7. Euphorbia hirsuta 2, Agrimonia eupatoria 1 in 8. Cyperus longus +, Lythrum junceum +, Mentha aquatica +, Verbena officinalis + in 9. Dipsacus fullonum +, Plantago major +, Polygonum lapathifolium +, Potentilla reptans +, Rumex crispus + in 10.

Localities: 1. Jaén: Santiago de la Espada, La Toba 30S WH3825, 100 m2. 2. Holotypus ass. Jaén: Santiago de la Espada, Huelga-Utrera. 30S WH 3623. 40 m2. 3. Jaén: Pontones, Puente río Segura. 30S WH 3020. 30 m2. 4. Albacete: Villaverde del Guadalimar, Fábrica de resinas. 30S WH 4157. 20 m2. 5. Jaén: Molinicos, Mesones. 30S WH 5660. 20 m2. 6. Albacete: Paterna del Madera, río Endrinales. 30S WH 5557. 20 m2. 7. Albacete: Paterna del Madera. 30 S WH 5768. 20 m2. 8. Albacete: Bogarra, El Batanero, 30 S WH 5970. 20 m2. 9. Jaén: Santiago de la Espada, Puente de Vites. 30 S WH 4623. 20 m2. 10. Jaén: Santiago de la Espada, Puente río Zumeta. 30 S WH 3815. 100 m2. 11. Synthesized table.

[Ríos & Alcaraz]

Sedetum campanulati ass. nova hoc loco (11.4.2)

(Myosotidion stoloniferae, Montio-Cardaminetalia, Montio-Cardaminetea)

Typus associatio: Table 87, rel. 3 [Avila: Solana de Béjar, Sierra de Béjar, Malillo creek. 30TTK6964. 1980 m, W, 1 m2].

Characteristic species: Sedum campanulatum (Sedum lagascae auct. non Pau).

Diagnosis: Association well characterized by the short-lived Sedum campanulatum, local endemic of the Bejaran-Gredensean Sector (Gredos, Tormantos and Béjar Mountains), that colonizes small temporary springs and rivulets among siliceous outcrops, flowing during thaw, in the supramediterranean and the lower orosubmediterranean belts (1300-2000 m).

Table 87

11.4.2 Sedetum campanulati
(Myosotidion stoloniferae, Montio-Cardaminetalia, Montio-Cardaminetea)

Altitude (1=10m)165175198180180
Number of species66977
Ordinal number123*45
Characteristic species:
Sedum campanulatum33524
Veronica langei1+1.3
Montia amporitana.+.+2
Epilobium anagallidifolium.11.2
Epilobium obscurum...11
Companion species:
Nardus stricta.+113
Juncus bulbosus+.+.2
Radiola linoides1..+2
Philonotis fontana..332
Viola palustris..122

Other species. Molineriella laevis 1, Leontodon carpetanus + in 1. Thymus pulegioides 2 in 2. Festuca rothmaleri 1, Carex iberica + in 3.

Localities: 1. Ávila: Sierra del Barco, Puerto Castilla, Garganta de la Vega. 30TTK7859. W, 1 m2. 2. Ávila: Guisando, Los Galayos, La Apretura. NW, 2 m2. 3. Holotypus ass. Ávila: Solana de Béjar, Sierra de Béjar, Malillo creek 30TTK6964. E, 1 m2. 4. Ávila: Hoyos del Espino, Sierra de Gredos, La Plataforma. 30TUK1060. N, 1 m2. 5. Synthesized table.

[Rivas-Martínez, Fernández-González, Sánchez-Mata & Sardinero]

Sedetum lagascae ass. nova hoc loco (9.4.9)

(Cicendion, Isoetetalia, Isoeto-Nanojuncetea)

Typus associatio: Table 88, rel. 4 [Avila: San Martín de la Vega del Alberche, Barbabúo creek. 30TUK1679. 1540 m, 1 m2].

Characteristic species: Sedum lagascae (Sedum maireanum).

Diagnosis: Association dominated by the succulent-leaved therophyte Sedum lagascae and several dwarf annual rushes (Juncus bufonius, J. tenageia, J. capitatus), growing on shallow mineral soils developed on siliceous rocks and submitted to short spring episodes of hydromorphy. They bloom in late spring-early summer and are mainly distributed in the meso and supramediterranean belts (up to 1800 m) of the Mediterranean West Iberian Province.

Remarks: Brullo & Minissale in Itinera Geobot. 11: 282. 1998 proposed the new association Junco bufonii-Sedetum lagascae based in the rel. 10, tb. 9 published by Sánchez-Mata (1989: 76) as "Comunidad de Juncus bufonius y Sedum lagascae". As the Italian authors do not give any comment in the text out of the number of the table, it seems possible to consider the association name as invalidly published (art. 2b), if contrarily we accept it as valid name the citation should be Junco bufonii-Sedetum lagascae Sánchez-Mata ex Brullo & Minissale 1998.

Table 88

9.4.9 Sedetum lagascae
(Cicendion, Isoetetalia, Isoeto-Nanojuncetea)

Altitude (1=10m)156118130154180110112124126136135
Number of species57861278913108
Ordinal number1234*567891011
Characteristic species:
Sedum lagascae3323343343V
Juncus bufonius2113133213V
Juncus capitatus.2.2..1.11III
Juncus tenageia...11..+3+III
Mentha pulegium....++..1.II
Radiola linoides..3....+..II
Ranunculus nodiflorus.....1+...II
Lythrum longidentatum......11..II
Scirpus setaceus........22II
Companion species:
Vulpia bromoides.+...1+1..III
Trifolium strictum.1...2..11III
Trifolium patens1.++......II
Molineriella laevis.1...1.+..II

Other species. Characteristic species: Juncus hybridus 1 in 10. Companion species: Aphanes microcarpa + in 1, 1 in 5. Anthoxanthum aristatum 2 in 3, + in 8. Sagina procumbens 1 in 3, + in 9. Ctenopsis delicatula + in 7 and 9. Euphrasia hirtella 1 in 9 and 10. Stellaria alsine 1 in 9 and 10. Trifolium dubium 1 in 1. Herniaria glabra + in 2. Trifolium arvense 1, Juncus articulatus + in 3. Aira caryophyllea 1 in 4. Campanula lusitanica 1, Moenchia erecta 1, Ranunculus paludosus 1, Trisetum ovatum 1, Bromus hordeaceus +, Myosotis persoonii +, Teesdalia coronopifolia + in 5. Vicia tetrasperma + in 7. Chamaemelum nobile + in 8. Prunella vulgaris 1, Ranunculus flammula + in 9. Cerastium vulgare + in 10.

Localities: 1. Ávila: San Martín de la Vega del Alberche, Barbabúo creek. 30TUK1679. 1 m2. 2. Madrid: El Paular. 30TVL2527. 10 m2. 3. Ávila: El Hornillo, Peña del Mediodía, Garganta del Arroyo de Aguas Frías. 30TUK1759. 1 m2. 4. Holotypus ass. Ávila: San Martín de la Vega del Alberche, Barbabúo creek. 30TUK1679. 1 m2. 5. Ávila: La Serrota, Cepeda la Mora, near of Risco del Cueto, Fuente Peguerinos. 30TUK2381. 1 m2. 6. Madrid: Gargantilla de Lozoya-Pinilla de Buitrago. 30TVL4036. 20 m2. 7. Madrid: Navarredonda-Pinilla de Buitrago. 30TVL4037. 5 m2. 8. Ávila: Navalperal de Tormes. 30TUK0569. 10 m2. 9. Ávila: San Martín del Pimpollar, Venta Rasquilla, Alberche river. 30TUK2871. 1 m2. 10. Ávila: San Martín del Pimpollar. 30TUK2570. 1 m2. 11. Synthesized table.

[Rivas-Martínez, Fernández-González, Sánchez-Mata & Sardinero]

Selaginello denticulatae-Saxifragetum gemmulosae F.J. Pérez, T.E. Díaz, P. Fernández & Salvo ass. nova hoc loco (30.5.3)

[Comunidad de Selaginella denticulata y Saxifraga gemmulosa F.J. Pérez, T.E. Díaz, P. Fernández & Salvo in Acta Bot. Malacitana 14: 177, tb. 5. 1989 (art. 2d)]
(Selaginello denticulatae-Anogrammion leptophyllae, Anomodonto-Polypodietalia, Anomodonto-Polypodietea)

Typus associatio: F.J. Pérez, T.E. Díaz, P. Fernández & Salvo in Acta Bot. Malacitana 14: 177, tb. 5 rel. 3. 1989.

Characteristic species: Saxifraga gemmulosa.

Diagnosis: Bryo-pteridophytic association with short thick leaved and pulviniform chamaephytes, characterized by the Bermejan serpentine endemic Saxifraga gemmulosa Boiss., typical of shady slopes and fissures of the thermo and mesomediterranean subhumid-humid belt of the Bermejan Subsector (Rondean Sector).

[Rivas-Martínez & Izco]

Senecioni laderoi-Juncetum inflexi ass. nova hoc loco (59.15.13)

[Senecioni laderoi-Juncetum inflexi M.E. García, L. Herrero, C. Pérez & Penas in M.E. García 1990 nom. inval. (art. 1)]
(Mentho-Juncion inflexi, Plantaginetalia majoris, Molinio-Arrhenatheretea)

Typus associatio: Table 89, rel. 4 [León: Villamanín. 30TTN85. 1140 m, NW, 5%, 20 m2].

Characteristic species (territorials): Carex binervis, Juncus acutiflorus, Juncus inflexus, Mentha longifolia, Senecio laderoi.

Diagnosis: Hygronitrophilous rush meadows growing on the banks of spring water courses in deep, damp and base-rich soils on the supratemperate belt of the Orocantabric Subprovince, mainly in the Ubinnean-Picoeuropean and Campurrian-Carrionese Sectors. This association is vicarious of the more acidophilous Mentho suaveolentis-Juncetum inflexi, from which it differs by the presence of Senecio laderoi and Cirsium pyrenaicum.

Table 89

59.15.13 Senecioni laderoi-Juncetum inflexi
(Mentho-Juncion inflexi, Plantaginetalia majoris, Molinio-Arrhenatheretea)

Altitude (1=10m)111144133114113114114116124
Number of species141414161718192117
Ordinal number1234*56789
Characteristic species:
Senecio laderoi21121222V
Juncus inflexus11223123V
Mentha longifolia13312333V
Trifolium pratense111+1.12V
Briza media11+..111IV
Prunella vulgaris.1.1111+IV
Juncus acutiflorus.11.1111IV
Holcus lanatus1..111.1III
Cirsium pyrenaicum1.3.22..III
Ranunculus repens.22..2.2III
Plantago lanceolata1+.1..1.III
Poa trivialis.1.+.11.III
Lathyrus pratensis.1..1.1.II
Achillea millefolium....1.+1II
Companion species:
Scrophularia alpestris.1.2.121III
Carex binervis.21..22.III
Rumex conglomeratus1...1..+II
Dipsacus fullonum1....+.1II

Other species. Characteristic species: Crepis capillaris 1 in 1 and 8. Lotus villosus + in 3, 3 in 4. Potentilla reptans + in 4 and 6. Agrostis capillaris 2 in 5 and 8. Dactylis glomerata 1 in 7 and 8. Festuca paniculata + in 2. Cynosurus cristatus 1, Senecio barbareifolius 1, Plantago major + in 3. Lychnis flos-cuculi 1, Arrhenatherum bulbosum +, Mentha xrotundifolia + in 4. Elymus caninus 1, Phleum bertolonii 1 in 5. Centaurea nigra + in 6. Trifolium repens 2, Bellis perennis 1 in 7. Anthoxanthum odoratum 1, Apium nodiflorum 1, Succisa pratensis + in 8. Companion species: Verbena officinalis + in 4, 1 in 6. Epilobium parviflorum 1 in 1 and 4. Galium verum + in 5, 1 in 7. Epilobium hirsutum 1 in 5 and 8. Dactylorhiza maculata 1 in 6 and 7. Scrophularia auriculata 2, Agrimonia eupatoria 1 in 1. Veronica beccabunga + in 3. Chaerophyllum hirsutum 1 in 5. Ononis spinosa + in 6. Iris germanica + in 7. Equisetum arvense + in 8.

Localities: 1. Palencia, Rebanal de las Llantas. 30TUN75. E, 2%, 15 m2. 2. León, Cubillas de Arbás, Collada de Alonga. 30TTN65. E, 10%, 10 m2. 3. Palencia, Ruesga, Brañosera. 30TUN74. N, 5%, 10 m2. 4. Holotypus ass. León, Villamanín. 30TTN85. NW, 5%, 20 m2. 5. Palencia, Traspeña de la Peña. 30TUN64. 25 m2. 6. León, Cármenes, Valverdín. 30TTN95. S, 10%, 20 m2. 7. León, Cubillas de Arbás, Collada de Alonga. 30TTN65. NE, 15%, 20 m2. 8. León, Sena de Luna. 30TTN55. N, 5%, 10 m2. 9. Synthesized table.

[M.E. García, L. Herrero, C. Pérez, Penas & F. Salegui]

Senecioni lopezii-Quercetum lusitanicae ass. nova hoc loco (75.11.3)

[Phillyreo angustifoliae-Quercetum fruticosae sensu auct iber. non Barbéro, Quézel & Rivas-Martínez in Phytocoenologia 9(3): 370, tb. 23. 1981]
(Quercion lusitanicae, Pistacio-Rhamnetalia alaterni, Querco-Fagetea)

Typus associatio: Cádiz: Sierra de Aljibe, upper Rocinejo River. 36º29'N-5º35'W. 780 m, NW, 10%, 40 m2. Characteristic species: 5 Quercus lusitanica, 2 Senecio lopezii, 1 Phillyrea angustifolia, 1 Serratula baetica subsp. alcalae, + Bupleurum foliosum, + Daphne gnidium, + Quercus suber (S2), + Rubia agostinhoi. Companion species: 2 Avenula lodunensis subsp. albinervis, 2 Caluna vulgaris, 1 Stauracanthus boivinii, + Agrostis curtisii, + Brachypodium gaditanum, + Halimium lasianthum, + Lavandula luisieri, + Satureja salzmannii, + Ulex borgiae.

Characteristic species (territorials): Avenula sulcata subsp. albinervis, Bupleurum foliosum, Quercus lusitanica, Senecio lopezii, Serratula alcalae, Ulex borgiae.

Diagnosis: Upper thermo-mesomediterranean humid-hyperhumid scrub and dwarf-scrub Aljibic endemic seral community, growing on poor silicic cambic and podzolic soils as a seral community of cork-oak forest (Teucrio baetici-Quercetum suberis). It could be considered as geovicariant of the Maghrebian-Tingitanian association Phillyreo angustifoliae-Quercetum (fruticosae) lusitanicae Barbero, Quézel & Rivas-Martínez in Phytocoenologia 9(3): 370, tb. 23. 1981; both association can be easily differenciated by: Senecio lopezii, Lavandula luisieri, Satureja salzmannii, Serratula alcalae and Ulex borgiae present in the Aljibic association, as well by Calicotome villosa, Erica arborea, Halimium halimifolium, Quercus coccifera and Teucrium fruticans very common in Morocco association but almost absent in the spanish geovicariant.

[Rivas-Martínez]

Sideritido ilicifoliae-Thymenion loscosii suball. nova hoc loco (64.5c)

[Rosmarino-Cistenion clusii Mateo in Publ. Ministerio Agricultura, Ser. Monogr. 31: 216. 1983 (art. 5)]
(Sideritido incanae-Salvion lavandulifoliae, Rosmarinetalia officinalis, Rosmarinetea officinalis)

Typus suballiancia: Sideritido ilicifoliae-Thymetum loscosii O. Bolòs & Molero Brion. in Molero Brion. in Butll. Inst. Catalana Hist. Nat., Sec. Bot. 51: 158, tb. 9. 1984.

Characteristic species: Centaurea linifolia subsp. linifolia, Euphorbia minuta subsp. minuta, Sideritis fruticulosa, Sideritis ilicifolia, Sideritis spinulosa, Teucrium gnaphalodes subsp. ilerdense, Thymus loscosii.

Diagnosis: Mesomediterranean and lower supramediterranean semiarid and dry semicontinental thyme and rosemary communities growing on clayey or calcareous eroded soils, generally with stones in surface (cobble and pebble sizes), spread in Low Aragonese Subprovince and adjacent Oroiberian territories.

Aphyllantho-Bupleuretum fruticescentis Br.-Bl. & O. Bolòs in Anales Estac. Exp. Aula Dei 5: 190, tb. 39. 1958 (64.5.21)

Distribution: Meso-lower supramediterranean dry Low Aragonese and adjacent territories.

Cytiso fontanesii-Cistetum clusii Br.-Bl. & O. Bolòs 1958 corr. O. Bolòs in Mem. Real Acad. Ci. Barcelona 38(1): 124. 1967 (64.5.22)

[Cytiso-Cistetum libanotidis Br.-Bl. & O. Bolòs in Anales Estac. Exp. Aula Dei 5: 165, tb. 35. 1958 (art. 43)]

Distribution: Lower mesomediterranean semiarid eastern Monegrensean Sector.

Rosmarino officinalis-Linetum suffruticosi Br.-Bl. & O. Bolòs in Anales Estac. Exp. Aula Dei 5: 142, tb. 30. 1958 (64.5.23)

Distribution: Mesomediterranean semiarid and dry Low Aragonese and adjacent territories.

Salvio lavandulifoliae-Ononidetum fruticosae Fernández-González, Loidi & A. Molina in Anales Jard. Bot. Madrid 42(2): 452, tb. 1. 1986 (64.5.24)

Distribution: Mesomediterranean dry Riojan-Estellan Sector.

Sideritido ilicifoliae-Thymetum loscosii O. Bolòs & Molero Brion. in Molero Brion. in Butll. Inst. Catalana Hist. Nat., Sec. Bot. 51: 158, tb. 9. 1984 (64.5.25)

Distribution: Lower mesomediterranean dry and semiarid Monegrensean Sector.

Sideritido spinulosae-Lavanduletum latifoliae A. Molina, Loidi & Fernández-González in Bot. Complut. 18: 31, tb. 5. 1993 (64.5.26)

Distribution: Meso-lower supramediterranean dry Sorian Oroiberian and Bardenan-Monegrensean Sectors.

[Rivas-Martínez, Cantó, Fernández-González & Sánchez-Mata]

Silybetum hispanici ass. nova hoc loco (34.11.7)

(Urtico piluliferae-Silybion mariani, Carthametalia lanati, Onopordenea acanthii, Artemisietea vulgaris)

Typus associatio: Table 90, rel. 1 [Ciudad Real: Pedro Muñoz, Laguna del Retamar. Disturbed places close to salt-marsh areas. 660 m, 10 m2].

Characteristic species: Silybum eburneum var. hispanicum, Silybum xgonzaloi.

Diagnosis: Community of annual or biennal great thistles, well characterized by the Iberian endemic Silybum eburneum var. hispanicum, geovicariant of the Mauritanous verticicolous also endemic Silybum eburneum var. eburneum, often accompanied by other great thistles such as Carthamus lanatus, Onopordum nervosum subsp. castellanum and Silybum marianum, with which can hybridize (Silybum xgonzaloi). These new Manchean-Murcian-Aragonian (Monegros) association grows on clayey or clayey-gypsicolous turned over soils with temporal hydromorphy and frequent vertic character.

Table 90

34.11.7 Silybetum hispanici
(Urtico piluliferae-Silybion mariani, Carthametalia lanati, Onopordenea acanthii, Artemisietea vulgaris)

Altitude (1=10m)66696870666668
Number of species10121816172015
Ordinal number1*234567
Characteristic species:
Silybum hispanicum334422V
Carthamus lanatus21++++V
Lactuca serriola122212V
Hirschfeldia incana1++.11V
Picnomon acarna2..+12IV
Carduus pycnocephalus.22.11IV
Silybum marianum+...33III
Centaurea calcitrapa.12..+III
Onopordum castellanum.+.++.III
Cichorium intybus..++.+III
Cirsium vulgare.+.1..II
Marrubium vulgare.+.1..II
Scolymus hispanicus..2+. II
Companion species:
Anacyclus clavatus1211.+V
Beta maritima.++.11IV
Bromus diandrus..1111IV
Hordeum leporinum.+..11III

Other species. Characteristic species: Carduus tenuiflorus +, Carlina hispanica +, Sylibum xgonzaloi + in 1. Onopordum castellanum x Onopordum acanthium 1, Onopordum acanthium + in 3. Chondrilla juncea 1 in 4. Companion species: Mantisalca salmantica 2 in 3, + in 4. Sonchus oleraceus 1 in 5 and 6. Avena barbata + in 5, 1 in 6. Papaver rhoeas + in 5, 1 in 6. Descurainia sophia + in 5 and 6. Phragmites communis + in 5 and 6. Sonchus xtoletanus + in 5 and 6. Hordeum leporinum x Hordeum marinum 2, Aeluropus littoralis + in 1. Medicago sativa +, Peganum harmala + in 3. Bromus hordeaceus 2, Verbena offcinalis 1, Gypsophila xcastellana +, Sonchus crassifolius + in 4. Althaea hirsuta + in 5. Hordeum marinum +, Podospermum laciniatum in 6.

Localities: 1. Holotypus ass. Ciudad Real: Pedro Muñoz, Laguna del Retamar. Disturbed places close to salt-marsh areas, 10 m2. 2. Cuenca: Las Mesas, "Laguna de Lara", 10 m2. 3. Toledo: Between Villacañas and Lillo, Hermitage of "Nuestra Señora de la Esperanza", 10 m2. 4. Toledo: Miguel Esteban, "Laguna de Paloma", Hermitage of "Nuestra Señora de Palomares", 10 m2. 5, 6. Ciudad Real: Pedro Muñoz, Laguna del Retamar. Disturbed places close to drainage channels grooved on old salt-marsh areas, 10 m2. 7. Syntesized table.

[Rivas-Martínez, Cantó, M.B. Crespo & Sánchez-Mata]

Sisymbrietum erysimoidis (Ladero, O. Socorro, Molero, M. López, Zafra, Marín, Hurtado & Pérez-Raya 1981) ass. nova, stat. nov. hoc loco (39.8.9)

[Basion.: Sisymbrio irionis-Malvetum parviflorae sisymbrietosum erysimoidis Ladero, O. Socorro, Molero, M. López, Zafra, Marín, Hurtado & Pérez-Raya in Anales Jard. Bot. Madrid 37(2): 739, tb. 2. 1981]
(Chenopodienion muralis, Chenopodion muralis, Chenopodietalia muralis, Stellarietea mediae)

Typus associatio: Ladero, O. Socorro, Molero, M. López, Zafra, Marín, Hurtado & Pérez-Raya in Anales Jard. Bot. Madrid 37(2): 739, tb. 2, rel. 4 (holotypus). 1981.

Characteristic species: Sisymbrium erysimoides.

Diagnosis: Thermo-lower mesomediterranean dry and semiarid, annual ephemeral ruderal, rural and urban early spring blooming and slightly shaded community, growing on Murcian-Almeriensian and Betic Provinces, and spread to Setabensean and Manchean biogeographic sector territories.

[Rivas-Martínez & Ladero]

Solenopsio balearicae-Naufragetum balearicae (Duvigneaud 1970) ass. nova, stat. nov. hoc loco (30.4.4)

[Sibthorpio africanae-Arenarietum balearicae adiantetosum capilli-veneris Duvigneaud in Bull. Soc. Roy. Bot. Belgique 76: 66. 1970 (basion.) (art. 27d, 46H)]
(Arenarion balearicae, Anomodonto-Polypodietalia, Anomodonto-Polypodietea)

Typus associatio: in Bull. Soc. Roy. Bot. Belgique 76: 66. 1970

Characteristic species: Naufraga balearica.

Diagnosis: Not arrived on time to be translated.

[Llorens & Gil]

Southbyo tophaceae-Pinguiculetum dertosensis ass. nova hoc loco (26.2.5)

(Pinguiculion longifoliae, Adiantetalia capilli-veneris, Adiantetea)

Typus associatio: Granada: Sierra Tejeda, Salto del Caballo. 30SVF0984. 1700 m, NW, 10 m2. Characteristic species: 2 Pinguicula dertosensis, 1 Eucladium verticillatum, 1 Southbya tophacea, + Pellia fabbroniana. Companion species: 1 Nostoc sp.

Characteristic species: Pinguicula dertosensis.

Diagnosis: Bryo-cormophyte community characterized by the allopoliploid endemic species Pinguicula dertosensis, locally growing on humid calcareous rocky banks and waterlogged pastures in meso and supramediterranean belts of Malacitan-Almijarensean Sector (Sierra Tejeda), Cazorlan territories (Cazorla Mountains) and Valencian-Tarraconensean Sector (Beceite Mountains).

[Asensi & Díez Garretas]

Spergulario rupicolae-Sedetum anglici ass. nova hoc loco (55.1.4)

(Sedion anglici, Sedo-Scleranthetalia, Sedo-Scleranthetea)

Typus associatio: Table 91, rel. 4 [Asturias: Gozón, Cabo Peñas. 30TTP3770. 100 m, 10%, 1 m²].

Characteristic species (territorials): Spergularia rupicola, Sedum anglicum.

Diagnosis: Coastal sparse grassland community of perennial grasses and small succulent chamaephytes, colonizing lithosols on slates, sandstones and quartzites. This association is characterized and well distinguished by Sedum anglicum and the halophilous species Spergularia rupicola and Festuca pruinosa. Its optimum occurs on the siliceous coastal capes of the thermotemperate humid belt in the Galician-Asturian Sector, although it also grows close to the heath communities of Angelico pachycarpae-Ulicetum maritimi and to the aerohaline grasslands of Crithmo-Armerion.

Table 91

55.1.4 Spergulario rupicolae-Sedetum anglici
(Sedion anglici, Sedo-Scleranthetalia, Sedo-Scleranthetea)

Altitude (1=10m)81010101010101010106628822107
Number of species101077967589679884558
Ordinal number1234*5678910111213141516171819
Characteristic species:
Sedum anglicum231333332321233433V
Spergularia rupicola (terr.)2132332..233212+11V
Festuca pruinosa (terr.).221+11211..+....+IV
Allium ericetorum+1++.....++1+.1...III
Koeleria albescens....1.......++121.II
Jasione gallaecica......++1+...+....II
Sedum hirsutum........+1........I
Companion species:
Hypochoeris radicata++11+11+11...31...IV
Dactylis maritima1+......+.+.++1+..III
Rumex biformis.++++++..+........III
Plantago coronopus++..1.....11.....1II
Scilla verna.1.1.....+...++...II
Armeria depilata..+.+.......+.+.+.II

Other species. Companion species: Lotus corniculatus + in 1, 14 and 17. Trifolium occidentale 1 in 5, 2 in 12 and 13. Desmazeria marina 3 in 11, 12 and 13. Sagina maritima 1 in 1, + in 2. Agrostis setacea 2 in 8, 1 in 9. Leucanthemum pluriflorum +, Poa annua 1 in 1. Lithodora prostrata + in 6. Ulex europaeus f. maritimus + in 7. Cirsium filipendulum + in 9. Anthyllis iberica +, Leontodon taraxacoides ssp. hispidus + in 12. Picris hieraciodes + in 18.

Localities: 1. Asturias: Cudillero, Cabo Vidio. 29TQJ3032. 15%, 4 m². 2. Asturias: Gozón, Cabo Peñas. 30TTP3770. 15%, 1 m². 3. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 4. Holotypus ass. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 5. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 6. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 7. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 2 m². 8. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 9. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 1 m². 10. Asturias: Gozón, Cabo Peñas. 30TTP3770. 10%, 10 m². 11. Asturias: Valdés, Cabo Busto. 29TQJ2704. 15%, 1 m². 12. Asturias: Valdés, Cabo Busto. 29TQJ2704. 15%, 2 m². 13. Asturias: Cudillero, Cala El Riego. 29TQJ2718. 15%, 3 m². 14. Asturias: Cudillero, Cabo Vidio. 29TQJ3032. 15%, 2 m². 15. Asturias: Cudillero, Cabo Vidio. 29TQJ3032. 20%, 2 m². 16. Asturias: Castropol, Penarronda. 29TPJ2462. 15%, 1 m². 17. Asturias: Castropol, Penarronda. 29TPJ2462. 15%, 1 m². 18. Asturias: Castrillón, Punta Vidrias. 29TQJ3040. 15%, 5 m². 19. Synthesized table.

[Arbesú, Bueno & F. Prieto]

Sporobolion arenarii (Géhu ex Géhu & Biondi 1994) all. nova, stat. nov. hoc loco (16.3)

[Sporobolenion arenarii Géhu ex Géhu & Biondi in Braun-Blanquetia 13: 14. 1994 (art. 27a, 46H)]
(Ammophiletalia, Ammophiletea)

Typus alliancia: Eryngio maritimi-Sporoboletum arenarii (Arènes ex Géhu & Biondi 1994) nom. nov. hoc loco.

Characteristic species: Sporobolus arenarius.

Diagnosis: Thermomediterranean semiarid and dry West Mediterranean pioneer perennial rizomatous grass communities, growing on first line of salt flat coastal dunes mostly of medium and coarse sand particles, characterized by Sporobolus arenarius.

Eryngio maritimi-Sporoboletum arenarii (Arènes ex Géhu & Biondi 1994) nom. nov. hoc loco (16.3.1)

[Group. à Sporobolus arenarius Arènes in Bull. Soc. Bot. France 71: 93. 1924 (art. 3c), Sporoboletum arenarii Arènes ex Br.-Bl., Roussine & Nègre, Group. Vég. France Médit.: 43. 1952 (pro syn.) non Rothmaler 1943 (art. 31), Sporoboletum arenarii Arènes ex Géhu & Biondi in Braun-Blanquetia 13: 12. 1994 non Rothmaler in Feddes Repert. Spec. Nov. Regni Veg. Beih. 128: 43. 1943 (art. 31) (nomencl. syn.), Sporobolo arenarii-Centaureetum maritimae Rivas Goday & Rigual 1959 (art. 37)]

Typus associatio: Géhu & Biondi in Braun-Blanquetia 13: 13, tb. 3, lectotypus hoc loco, rel. 7. 1994 [Corse: Etang d"Arasu, 10 m2]. Caracteristic species: 4 Sporobolus arenarius, + Eryngium maritimum. Companion species: + Salsola kali.

Characteristic species (territorial): Sporobolus arenarius.

Diagnosis: Thermomediterranean dry pioneer community dominated by Sporobolus arenarius growing on coastal salt flat dunes of medium and coarse sand particles, spread in Tyrrhenian and West European Mediterranean drainage-bassin.

Sporoboletum arenarii Rothmaler in Feddes Repert. Spec. Nov. Regni Veg. Beih. 128: 43. 1943 (16.3.2) (lectotypus hoc loco, l.c. rel. 291)

Distribution: Lusitanian-Andalusian coast.

[Rivas-Martínez & Cantó]

Stipetum cazorlensis (J. Torres & Cano in J. Torres, A. García, Salazar & Cano 2001) ass. nova, stat. nov. hoc loco (56.5.5)

[Helictotricho filifolii-Festucetum scariosae stipetosum cazorlensis J. Torres & Cano in J. Torres, A. García, Salazar & Cano in Phytocoenologia 31(1): 136, tb. 2. 2001 (basion.) (art. 27d, 46H)]
(Stipion parviflorae, Lygeo-Stipetalia, Lygeo-Stipetea)

Typus associatio: Helictotricho filifolii-Festucetum scariosae stipetosum cazorlensis J. Torres & Cano in J. Torres, A. García, Salazar & Cano in Phytocoenologia 31(1): 136, tb. 2, rel. 3. 2001 Holotypus, Jaén: Arroyo de los Fresnos, near Cortijo Palomares (Campillo de Arenas) 30SVG3967, alt.: 1110 m, 30 m2]. Characteristic species: 4 Stipa cazorlensis, 1 Arrhenatherum album, 1 Avenula gervaisii, 1 Dactylis hispanica. Companion species: 1 Aegilops triuncialis, 1 Avena barbata, 1 Medicago sativa, 1 Plantago lanceolata, + Convolvulus arvensis, + Leontodon longirostris, + Silene vulgaris, + Trifolium stellatum.

Characteristic species: Stipa bufensis, Stipa cazorlensis.

Diagnosis: Meso-supramediterranean dry and lower subhumid somewhat open slightly subnitrophilous pioneer perennial grassland community, characterized by Stipa cazorlensis, Stipa bufensis and Stipa iberica subsp. austroiberica, growing on carbonated soils in Subbetic and Guadician-Bacensean Sectors.

[Rivas-Martínez]

Suaedetum spicato-splendentis ass. nova hoc loco (25.1.6)

(Thero-Suaedion, Thero-Suaedetalia, Thero-Salicornietea)

Typus associatio: Table 93, rel. 6 [Ciudad Real: Alcázar de San Juan, La Laguna. 30VJ791605. 680 m, 1 m2]

Characteristic species (territorials): Suaeda spicata, Suaeda splendens.

Diagnosis: Mesomediterranean and lower supramediterranean dry semicontinental pioneer temporary inundated halo-nitrophilous salt-pans community of therophytic succulent plants, spread in Mediterranean Central Iberian Province mostly on continental drainage-bassin of Manchean Sector pools, characterized by Suaeda spicata and Suaeda splendens.

Table 93

25.1.6 Suaedetum spicato-splendentis
(Thero-Suaedion, Thero-Suaedetalia, Thero-Salicornietea)

Altitude (1=10m)7266676772686668727069
Number of species45556666696
Ordinal number123456*7891011
Characteristic species:
Suaeda spicata5355441444V
Suaeda splendens+3...142+1IV
Salicornia patula..1+++.+..III
Cressa cretica.+..+.....I
Beta maritima.+........I
Companion species:
Frankenia pulverulenta1.+2.2+++.IV
Sphenopus divaricatus..+++.+.++IV
Hordeum marinum..1.++..+1III
Parapholis incurva.....+.+.1II

Other species. Companion species: Polypogon maritimus + in 1 and 5. Atriplex prostrata 2 in 2 and 10. Bassia hyssopifolia 1 in 7 and 10. Puccinellia caespitosa + in 8 and 9. Lolium rigidum + in 4. Aeluropus littoralis + in 7. Elymus curvifolius +, Spergularia marina + in 10.

Localities: 1. Toledo: La Puebla de Almoradiel. Hermitage of "Nuestra Señora de Palomares", "Lagunas de Paloma". 30SVJ848766. 4 m2. 2. Ciudad Real: Pedro Muñoz. "Laguna del Pueblo". 30SWJ047634. 4 m2. 3. Ciudad Real: Alcázar de S. Juan. "Laguna de las Yegüas". 30SVJ757627. 4 m2. 4. Ciudad Real: Alcázar de S. Juan. "Laguna de las Yegüas". 30SVJ757626. 2 m2. 5. Toledo: La Puebla de Almoradiel. Hermitage of "Nuestra Señora de Palomares", "Lagunas de Paloma". 30SVJ849765. 2 m2. 6. Holotypus ass. Ciudad Real: Alcázar de S. Juan. "La Laguna". 30SVJ791605. 1 m2. 7. Ciudad Real: Pedro Muñoz. Hermitage of "S. Cristóbal", "Laguna de Navalafuente", nitrophilous plant community stayed in old-crops areas. 30SWJ063638. 4 m2. 8. Toledo: Villafranca de los Caballeros. "Lagunilla de la Sal". 30SVJ714657. 2 m2. 9. Toledo: La Puebla de Almoradiel. Hermitage of "Nuestra Señora de Palomares", "Lagunas de Paloma". 30SVJ849766. 4 m2. 10. Toledo: Lillo, Hermitage of "Nuestra Señora de la Esperanza", salt-marsh area with old crops now abandoned. 30SVJ714906. 4 m2. 11. Synthesized table.

[Rivas-Martínez, Cantó & Sánchez-Mata]

Suaedo braun-blanquetii-Tamaricetum canariensis ass. nova hoc loco (70.3.6)

[Agrostio-Tamaricetum canariensis inuletosum crithmoidis Fernández-González, A. Molina & Loidi in Acta Bot. Malacitana 15: 314, tb. 2. 1990 (corresp. name)]
(Tamaricion boveano-canariensis, Tamaricetalia, Nerio-Tamaricetea)

Typus alliancia: Table 92, rel. 4 [Zaragoza: Tauste. Salt-marsh areas at Baranco de Valdespartera, between Tauste and Castejón de Valdejasa. 30TXM508480. 280 m, 100 m2].

Characteristic species (territorials): Arthrocnemum macrostachyum, Elytrigia curvifolia, Inula crithmoides, Limonium ruizii, Sarcocornia alpini, Suaeda braun-blanquetii, Tamarix canariensis.

Diagnosis: Middle basin of the Ebro river halo-hydromorphic microforests in which Tamarix canariensis is dominant. These salt marsh Tamarix communities, from pond borders, endorreic depressions, intermittent water courses, etc., represent the head of the series or potential natural vegetation on saline hydromorphic soils, in the mesomediterranean xeric or pluviseasonal dry belts of the Bardenan-Monegrensean Sector, although its presence could also be interpreted in the Manchean Sector. A great number of halophilous plants grow in its under-storey or among the Tamarix and belong to the classis Sarcocornietea fruticosae such as: Arthrocnemum macrostachyum, Inula crithmoides, Limonium ruizii, Sarcocornia alpini, Suaeda braun-blanquetii, etc. These habitats, that are given priority by the European Union Directives, are greatly threatened because of the drainage and the agricultural development of irrigated land.

Table 92

70.3.6 Suaedo braun-blanquetii-Tamaricetum canariensis
(Tamaricion boveano-canariensis, Tamaricetalia, Nerio-Tamaricetea)

Altitude (1=10m)39262628262529
Number of species56679-7
Ordinal number1234*567
Characteristic species:
Tamarix canariensis55355VV
Companion species:
Suaeda braun-blanquetii31+23VV
Inula crithmoides++.2+IIIIII
Phragmites communis+++.+IIIIV
Juncus maritimus.332.IIII
Elytrigia curvifolia.++.2.III
Atriplex halimus2...1IVII
Limonium ruizii..++..II
Arthocnemum macrostachyum...+.III
Sarcocornia alpini...1..I
Puccinellia caespitosa....+III

Other species. Companion species: Allium roseum +, Dorycnium pentaphyllum + in 5.

Localities: 1. Huesca: Senés de Alcubierre. Balsa Buena, salt-marsh areas, 30TYM085424, 100 m2. 2, 3. Huesca: Sariñena. Salt-marsh areas between Sariñena and Pallaruelo de Monegros, 30TYM336244, 80 m2. 4. Holotypus ass. Zaragoza: Tauste. Salt-marsh areas at Barranco de Valdespartera between Tauste and Castejón de Valdejasa, 30TXM508480, 100 m2. 5. Huesca: Sariñena. Salt-marsh areas at Barranco de S. Juan near S. Juan de Flumen, 30TYM310280, 100 m2. 6. Fernández-González, Molina & Loidi in Acta Bot. Malacitana 15: 314, tb. 2, 13 rel. 1990 (sub Agrostio stoloniferae-Tamaricetum canariensis tamaricetosum canariensis et inuletosum crithmoidis). 7. Synthesized table.

[Rivas-Martínez, Cantó & Sánchez-Mata]